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Whitehead, Wayne F.

Department: 
Agricultural Research
Location: 
Stallworth Building
Phone: 
(478) 825-6808
Email: 
whiteheadw@fvsu.edu
Fax: 
(478) 825-6376

1990-present: Research Professional, Agronomist/Production Specialist - Fort Valley State University

1989-1990: Research Associate, Agronomist - University of Tennessee, Highland Rim Experiment Station

1988-1989: Postdoctoral Research Associate - University of Tennessee-Knoxville

1984-1988: Graduate Research Assistant - University of Tennessee-Knoxville

1981-1983: Graduate Research Assistant - Alabama A&M University

1978-1981: Undergraduate Research Assistant - Alabama A&M University

1972-1976: U.S. Naval Hospital Corpsman, Head Hospital Corpsman - Guam Naval Hospital

Faculty Bio
Education: 

Doctor of Philosophy: Plant and Soil Science - The University of Tennessee-Knoxville (1988)

Master of Science: Plant and Soil Science - Alabama Agricultural & Mechanical University (1984)

Bachelor of Science (Honors): Horticulture - Alabama Agricultural & Mechanical University (1981)

Publications: 

Peer-Review Publications:
1.Sainju, U. M., H. H. Schomberg, B. P. Singh, W. F. Whitehead, P. G. Tillman and S. L. Lachnicht-Weyers. 2007. Cover crop effect on soil carbon fractions under conservation tillage cotton . Soil & Tillage Res. 63: 167-179.
2.Sainju, U.M., B.P. Singh, W.F. Whitehead and S. Wang. 2007. Accumulation and crop uptake of soil mineral nitrogen as influenced by tillage, cover crops, and nitrogen fertilization. Agron. J. 99:682-691.
3.Sainju, U.M., B.P. Singh, W.F. Whitehead, and S. Wang. 2006. Carbon supply and storage in tilled and nontilled soils as influenced by cover crops and nitrogen fertilization. J. Environ. Qual. 35:1507-1517.
4.Sainju, U.M., W.F. Whitehead, B.P. Singh, and S. Wang. 2006. Tillage, cover crops, and nitrogen fertilization effects on soil nitrogen and cotton and sorghum yields. Europ. J. Agron. 25:372-382.
5.Sainju, U. M., W. F. Whitehead and B. P. Singh. 2005. Biculture legume-cereal cover crops for enhanced biomass yield and carbon and nitrogen. Agron. J. 97: 1403-1412.
6.Sainju, U. M., B. P. Singh and W. F. Whitehead. 2005. Tillage, cover crops, and nitrogen fertilization effects on cotton and sorghum root biomass, carbon, and nitrogen. Agron. J. 97: 1279-1290.
7.Sainju, U. M., W. F. Whitehead and B. P. Singh. 2005. Carbon accumulation in cotton, sorghum, andunderlying soil as influenced by tillage, cover crops, and nitrogen fertilization. Plant and Soil 273: 219-234.
8.Whitehead, W. F. and B. P. Singh. 2005. Fresh market tomato gas exchange, biomass, and fruit yield are similar with legume cover crops or synthetic nitrogen fertilizer . HortScience 40(1):209-213.
9.Terrill, T. H., W. F. Whitehead, R. G. Durham, C. S. Hoveland, B. P. Singh and S. Gelaye. 2004. Preference of grazing goats for cool-season annual clovers. South African Journal of Animal Science 34: 107-109.
10.Sainju, U. M., W. F. Whitehead and B. P. Singh . 2003. Agricultural management practices to sustain crop yields and improve soil and environmental qualities. The Scientific World J. 3: 768-789.
11.Sainju, U. M., W. F. Whitehead and B. P. Singh . 2003. Cover crop and nitrogen fertilization effects on soil aggregation and carbon and nitrogen pools. Can. J. Soil Sci. 83: 155-165.
12.Whitehead, W. F., J. Carter and B. P. Singh. 2002. Effect of planting date on vegetable amaranth leaf yield, plant height and gas exchange. HortScience 37(5):773-777.
13.Sainju, U. M., B. P. Singh and W. F. Whitehead. 2002. Long-term effects of tillage, cover crops, and nitrogen fertilization on organic carbon and nitrogen concentrations in sandy loam soils in Georgia. Soil & Tillage Res. 63: 167-179.
14.Sainju, U. M., B. P. Singh and W. F. Whitehead. 2001. Comparison of the effects of cover crops and nitrogen fertilization on tomato yield, root growth, and soil properties. Scientia Horticulturae 91: 201-214.
15.Whitehead, W. F., and B. P. Singh. 2000. Yield, time of maximum CO2 exchange rate and leaf-area index of >Clemson Spineless= Okra are affected by within-row spacing. HortScience 35(5):849-852.
16.Sainju, U. M., B. P. Singh and W. F. Whitehead. 2000. Cover crop and nitrogen fertilization effects on soil carbon and nitrogen and tomato yield. Can. J. Soil Sci. 80: 523-532.
17.Singh, B. P. and W. F. Whitehead. 1998. Production Potential of Pointed Gourd in Temperate Climates, p. 397-399. In J. Janick (ed.) Perspectives on New Crops and New Uses.
18.Sainju, U. M., B. P. Singh and W. F. Whitehead. 1998. Cover crop root distribution and its effects on soil nitrogen cycling. Agron. J. 90: 511-518.
19.Singh, B. P. and W. F. Whitehead. 1996. Management methods for producing vegetable amaranth, p.511- 515. In J. Janick (ed.). Progress in New Crops.
20.Carter, Johnny, B. P. Singh and W. F. Whitehead. 1996. Dikegulac, but not benzyladenine enhances the aesthetic quality of Boston fern. HortScience 31(6):978-980.
21.Whitehead, W. F. and B. P. Singh. 1995. Leaf age affects gas exchange in okra. HortScience 30:1017-1019. 22) Singh, B. P. and W. F. Whitehead. 1993. Population density and soil pH effects on vegetable amaranth
22.production, p.562-564. In J. Janick and J. E. Simon (eds.). New Crops. 23) Singh, B. P. and W. F. Whitehead. 1992. Response of vegetable amaranth to differing soil pH and moisture regimes. Acta Horticulturae. 318:225-229.
23.Whitehead, W. F. and F. L. Allen. Fall, 1992. Comparison of Selection Intensities for Effectiveness of Selecting Superior Soybean Lines in High-versus Low-Stress Environments. Tennessee Farm and Home Science. 164:10-16.
24.Singh, B. P., K. A. Tucker, J. D. Sutton, H. L. Bhardwaj and W. F. Whitehead. 1991. Flooding reduces gas exchange and growth in snap bean. HortSci. 26:372-373.
25.Whitehead, W. F. and F. L. Allen. 1990. High- versus low-stress yield test nursery environments for selecting superior soybean lines. Crop Science. 30:912-918.
26.Whitehead, W. F., V. T. Sapra and D. R. Rao. 1989. Sulfur-containing amino acids and protein content in selected soybean progenies. Soybean Genetics Newsletter. 16:74-80.

Proceedings:
1.H. P. Singh, B. P. Singh, W. F. Whitehead and J. P. Wilson. 2011. Determining the genetic divergence of recombinant indred line(RIL) segregates of pearl millet for photosynthetic and water use efficiency traits. In Proceedings of the 16th Biennial Research Symposium of the Association of Research Directors, Inc., Atlanta, GA, April9-13 p. 193-194.
2.B. P. Singh, Whitehead, W. F., Terrill, T. H. and Singh, H. P.. 2010. Evaluation of napiergrass for dual use as an animal fodder and biofuel feedstock. Proceedings of the 19th Annual Meeting of the American Forage and Grassland Council, Brazil. (in print)
3.Terrill, T. H., W. R. Getz, S. L. Dzimianski, W. F. Whitehead, B. P. Singh and J. P. Muir. 2009. Effect of repeated or one-time grazing of kudzu by sheep and goats in central Georgia. 6 pp. Proceedings of the 18th Annual Meeting of the American Forage and Grassland Council, June 21-24, Grand Rapids MI. AFGC, on CD.
4.Gulia S. K., W. F. Whitehead B. P. Singh and J. P. Wilson. 2009. Residue amount, type, and placement effects on soil organic carbon and total nitrogen. In Proceedings of the 15th Biennial Research Symposium of the Association of Research Directors, Inc., Atlanta, GA, March 28-April 1. p. 117-118.
5.Terrill, T. H., W. F. Whitehead, K. H. Quesenberry, P. Mislevy, A. Blount, M. J. Williams, B. P. Singh and S. Gelaye. Growth and survival of rhizoma peanut in central Georgia. Proceedings of the American Forage and Grassland Council, Vol 12. April 2003, Lafeyette, Louisiana.06) Singh, B. P., U. M. Sainju, D. Lee, and W. F. Whitehead. 2003. Characteristics of light texture soils under high resdue conservation tillage systems for corn. Soil Tillage Research Organization 16th. Triennial Conference, Brisbane, Australia, 14-18 July, 2003. p. 139.
6.Sainju, U. M., W. F. Whitehead and B. P. Singh. 2003. Residue amount, type, and placement effects on soil organic carbon and total nitrogen. In Proceedings of the 13th Biennial Research Symposium of the Association of Research Directors, Inc., Atlanta, GA, 29 March-April. p. 112.
7.Terrill, T. H., W. F. Whitehead, R. G. Durham, C. S. Hoveland, B. P. Singh and S. Gelaye. Preference of grazing goats for cool-season annual clovers. Proceedings of the XI American Forage and Grassland Council. July 2002, Bloomington, Minnesota.
8.Whitehead, W. F., T. H. Terrill, B. P. Singh and S. Gelaye. Amaranth productivity and nutrient composition in central Georgia. Proceeding of the XIX International Grassland Congress. February 2001, Piracicaba, Brazil.
9.Phillips, D., U. M. Sainju, B. P. Singh, and W. F. Whitehead. 2000. Tillage and cover crop effects on soil properties and corn yield. In Proceedings of the 12th Biennial Research Symposium of the Association of Research Directors, Inc., Washington, D. C.

Abstracts and Presentations:
1.H. P. Singh, B. P. Singh, W. F. Whitehead and J. P. Wilson. 2011. Determining the genetic divergence of recombinant indred line(RIL) segregates of pearl millet for photosynthetic and water use efficiency traits. The 16th Biennial Research Symposium of the Association of Research Directors, Inc., Atlanta, GA, April9-13 p. 193-194.
2.B. P. Singh, Whitehead, W. F., Terrill, T. H. and Singh, H. P.. 2010. Evaluation of napiergrass for dual use as an animal fodder and biofuel feedstock. The 19th Annual Meeting of the American Forage and Grassland Council, Brazil.
3.W. F. Whitehead and B. P. Singh. 2010. Influence of Nitrogen Fertilization Rates and Legume Cover Crop on Water Use and Yield Response of Two Sweet Corn Cultivars. Annual ASHS Meetings.
4.W. F. Whitehead and B. P. Singh. 2010. Impact of Inorganic Nitrogen and Legume-Non Legume Cover Crops on Above Ground Biomass Yields and Leaf Area Index of Two Sweet Corn Cultivars. Annual ASHS Meetings.
5.W. F. Whitehead and B. P. Singh. 2009. Tillage Impact on Water Use and Yield of Three Sweet Corn Cultivars. Annual ASHS Meetings.
6.W. F. Whitehead and B. P. Singh. 2009. Influence of Tillage on Leaf Area Index and Above Ground Biomass Yields of Six Sweet Corn Cultivars. Annual ASHS Meetings.
7.W. F. Whitehead and B. P. Singh. 2008. Comparison of Cover Crops and Inorganic Nitrogen Effect on Leaf Area Index and Above Ground Biomass Yields of Southern Pea and Common Dry Bean. Annual ASHS Meetings.
8.W. F. Whitehead and B. P. Singh. 2008. Cereal Cover Crop vs. Synthetic Nitrogen Effect on Water Use and Yield Response of Southern Pea and Common Dry Bean. Annual ASHS Meetings.
9.W. F. Whitehead and B. P. Singh. 2007. Water Use and Yield Response of Three Common Dry Bean Cultivars to Cereal Cover Crop and Tillage Practices. Annual ASHS Meetings.
10.W. F. Whitehead and B. P. Singh. 2007. Cover Crop and Tillage Effects on Leaf Area Index and Above Ground Biomass Yields of Six Common Dry Bean Cultivars. Annual ASHS Meetings.
11.W. F. Whitehead and B. P. Singh. 2006. Effect of Biculture Legume-Cereal Cover Crops on Kernel Weight and Yields of Sweet Corn grown With or Without Synthetic Nitrogen. Annual ASHS Meetings.
12.W. F. Whitehead and B. P. Singh. 2006. Influence of Single and Biculture Cover Crops With or Without Synthetic Nitrogen on Leaf Area Index and Above Ground Biomass Yields of Sweet Corn. Annual ASHS Meetings.
13.W. F. Whitehead and B. P. Singh. 2005. Effect of a Non-leguminous Cover Crop, Tillage Practices, and Inorganic Nitrogen on Above Ground Biomass Production and Yields of Southernpea. Annual ASHS Meetings.
14.W. F. Whitehead and B. P. Singh. 2005 Influence of Single or Mixed Cover Crops on Biomass Production and Fruit Yields of Eggplant grown With or Without Synthetic Nitrogen. Annual ASHS Meetings.
15.Sainju, U. M., W. F. Whitehead and B. P. Singh . 2004. Tillage, cover crops and nitrogen fertilization effects on cotton and sorghum yields and nitrogen uptake. Annual ASA-CSSA-SSA Meetings. CD (Abstr.).
16.Sainju, U. M., W. F. Whitehead and B. P. Singh . 2004. Tillage, cover crops, and nitrogen fertilization effects on cotton and sorghum yields and nitrogen uptake. Annual ASA-CSSA-SSA Meetings.
17.W. F. Whitehead and B. P. Singh. 2004. Comparison of cover crops alone or in mixtures with inorganic nitrogen for above ground biomass production and fruit yields in bell pepper. Annual ASHS Meetings.
18.W. F. Whitehead and B. P. Singh. 2004. Cover crops their mixture and inorganic nitrogen effect on bell pepper gas exchange at three stages of fruiting. Annual ASHS Meetings.
19.Sainju, U. M., W. F. Whitehead and B. P. Singh . 2003. Carbon and nitrogen inputs from cover crops for soil sequestration. Annual ASA-CSSA-SSA Meetings.
20.W. F. Whitehead and B. P. Singh. 2003. Cover crop mixtures & Components vs. synthetic nitrogen effect on above ground biomass and yields of tomato. Annual ASHS Meetings.
21.W. F. Whitehead and B. P. Singh. 2003. Organic vs. inorganic nitrogen effect on tomato gas exchange at three stages of fruiting. Annual ASHS Meetings.
22.Singh, B. P., U. M. Sainju, D. Lee, and W. F. Whitehead. 2003. Characteristics of light texture soils under high resdue conservation tillage systems for corn. Soil Tillage Research Organization 16th. Triennial Conference, Brisbane, Australia, 14-18 July, 2003. p. 139.23) Sainju, U. M., W. F. Whitehead and B. P. Singh. 2003. Residue amount, type, and placement effects on soil organic carbon and total nitrogen. In Proceedings of the 13th Biennial Research Symposium of the Association of Research Directors, Inc., Atlanta, GA, 29 March-April. p. 112.
23.Terrill, T. H., W. F. Whitehead, R. G. Durham, C. S. Hoveland, B. P. Singh, and S. Gelaye. 2003. Preference of grazing Goats for cool-season Annual clovers. In Proceedings of the 13th Biennial Research Symposium of the Association of Research Directors, Inc., Atlanta, GA.
24.J. Carter, G. J. Johnson and W. F. Whitehead,. 2002. Effect of different concentrations of rooting hormones on the propagation of two cultivars of daylily from proliferations. 25th International Horticultural Congress.
25.W. F. Whitehead and B. P. Singh. 2002. Winter cover vs. synthetic N effect on yield, plant dry matter and photosynthesis of bell pepper. 25th International Horticultural Congress.
26.W. F. Whitehead, J. Carter and B. P. Singh. 2002. Effect of planting date on vegetable amaranth leaf yield, plant height and gas exchange. 25th International Horticultural Congress.
27.B. P. Singh and W. F. Whitehead. 2002. Pointed gourd (Trichosanthes dioica): A choice vegetable of India can be grown successfully in the southern United States. 25th International Horticultural Congress.
28.W. F. Whitehead and B. P. Singh. 2001. Organic vs. inorganic nitrogen effect on vegetative matter and yields of eggplant. Annual ASHS Meetings.
29.W. F. Whitehead and B. P. Singh. 2001. Eggplant gas exchange response to nitrogen sources at three stages of fruiting. Annual ASHS Meetings.
30.Phillips, D., U. M. Sainju, B. P. Singh, and W. F. Whitehead. 2000. Tillage and cover crop effects on soil properties and corn yield. In Proceedings of the 12th Biennial Research Symposium of the Association of Research Directors, Inc., Washington, D. C.
31.Sainju, U. M., B. P. Singh, and W. F. Whitehead. 2000. Soil carbon sequestration and nitrogen management with cover crops and nitrogen fertilization. Soil Science Society of America Symposium.
32.W. F. Whitehead and B. P. Singh. 2000. Three year effects of nitrogen sources on plant component and yields of tomato. Annual ASHS Meetings.
33.W. F. Whitehead and B. P. Singh. 2000. Three years of tomato gas exchange response to nitrogen sources at fruiting. Annual ASHS Meetings.
34.W. F. Whitehead and B. P. Singh. 1999. Influence of tillage and N source on gas exchange in tomato. Annual ASHS Meetings.
35.B. P. Singh and W. F. Whitehead. 1999. Tomato flowering and fruiting under reduced tillage. Annual ASHS Meetings.
36.W. F. Whitehead and B. P. Singh. 1998. Yield and fruiting span of tropical vegetable crop parwal in a temperate climate. Annual ASHS Meetings.
37.W. F. Whitehead and B. P. Singh. 1998. Tomato growth and yield response to cover crop and fertilizer nitrogen. Annual ASHS Meetings.
38.B. P. Singh, U. M. Sainju, and W. F. Whitehead. 1998. Nitrogen assimilation and biomass yield of winter cover crops used in sustainable horticultural crop production. Annual ASHS Meetings.
39.W. F. Whitehead and B. P. Singh. 1997. Introduction of high value Asian vegetables in Georgia. Georgia's Annual Ag. Exposition and Sunbelt Trade.
40.W. F. Whitehead and B. P. Singh. 1997. Gas exchange behavior of male and female Parwal plants. Annual ASHS Meetings.
41.W. F. Whitehead and B. P. Singh. 1997. Effect of N source on photosynthesis and plant dry matter yield of tomato. Annual ASHS Meetings.
42.B. P. Singh and W. F. Whitehead. 1997. Winter legume Cover for sustainable tomato production. Annual ASHS Meetings.
43.J. Carter, B. P. Singh and W. F. Whitehead. 1997. Growth and development response of flowering cabbage and kale cultivars to four growth retardents. Annual ASHS Meetings.
44.W. F. Whitehead and B. P. Singh. 1996. Gas exchange in tomato in a cover crop-tomato cropping system. Annual ASHS Meetings.
45.W. F. Whitehead and B. P. Singh. 1995. The impact of cover crops for vegetable production. Georgia's Vegetable and Small Fruit Conference and Trade Show.
46.B. P. Singh and W. F. Whitehead. 1995. Performance of Bittermelon in temperate climate. Annual ASHS Meetings.
47.W. F. Whitehead and B. P. Singh. 1995. Photosynthetic rates and yield components of eight vegetable amaranthus genotypes. Annual ASHS Southern Regional Meetings.
48.B. P. Singh and W. F. Whitehead. 1994. Soil moisture requirements for optimal growth of vegetable amaranth. Annual ASHS Meetings.
49.W. F. Whitehead and B. P. Singh. 1994. Vegetative growth of vegetable amaranth as affected by nitrogen fertilization. Annual ASHS Southern Regional Meetings.
50.W. F. Whitehead and B. P. Singh. 1993. Effect of nitrogen fertilization on growth and yield of amaranth. Annual ASHS Meetings.
51.W. F. Whitehead and B. P. Singh. 1993. Bitter Gourd: A potential crop for small farms. Annual ASHS Meetings.
52.B. P. Singh and W. F. Whitehead. 1993. Determination of optimal span on vegetable amaranth production. Annual ASHS Meetings.
53.B. P. Singh and W. F. Whitehead. 1992. Soil pH and moisture effects on vegetable amaranth production. Annual ASHS Meetings.
54.W. F. Whitehead and B. P. Singh. 1992. Vegetable amaranth performance at different soil moisture levels. Annual ASHS Southern Regional Meetings.
55.B. P. Singh and W. F. Whitehead. 1991. Population density and soil pH effects on vegetable amaranth production. 2nd Nat. Symposium, New Crops- Exploration, Research, Commercialization.
56.W. F. Whitehead and B. P. Singh. 1991. Gas exchange, chlorophyll content and leaf area index of okra at six in-row plant densities. Annual ASHS Meetings.
57.W. F. Whitehead and B. P. Singh. 1991. Diurnal and leaf age effects on gas exchange in okra. Annual ASHS Meetings.
58.B. P. Singh and W. F. Whitehead. 1991. Performance of Amaranth at six in-row plant densities. Annual ASHS Meetings.
59.V. T. Sapra, W. F. Whitehead, D. R. Rao, and A. Hassanalizadeh. 1988. Genetic variation for sulfur containing amino acids and protein content in selected soybean progenies. 16th International Congress of Genetics.55) W. F. Whitehead and F. L. Allen. 1987. High- versus low-stress yield test nursery environments for selecting superior soybean lines. Annual ASA-CSSA-SSA Meetings.
60.W. F. Whitehead, F. L. Allen and D. A. Lietzke. 1986. Effects of soil properties on root growth and seed yield of soybean lines: I. Chemical Properties. Annual ASA-CSSA-SSA Meetings.
61.D. A. Lietzke, W. F. Whitehead and F. L. Allen. 1986. Effects of soil properties on root growth and seed yield of soybean lines: II. Physical Properties. Annual ASA-CSSA-SSA Meetings.
62.W. F. Whitehead, V. T. Sapra, D. R. Rao and C. B. Chawan. 1983. Evaluation of soybean crosses based on sulfur-containing amino acids to protein content level. Annual ASA-CSSA-SSA Meetings.
63.W. F. Whitehead and V. T. Sapra. 1983. Genetic variability for protein content and agronomic characters in selected soybean crosses. Annual Southern ASA-CSSA-SSA Regional Meetings.
64.W. F. Whitehead, T. Mebrahtu and V. T. Sapra. 1980.Effects of seed size on germination and growth yield characteristics of three soybean cultivars. ASA-CSSA-SSA Annual Meetings.

Research Interests: 

Utilization of Agronomy/Horticultural Principles, Soil Science, Physiology, Breeding/Propagation Methods and
Experimental Design for the improvement of yield, nutrition, adaptability and utilization (animal fodder/biofuel feedstock) of plants (AGRONOMIC, HORTICULTURAL, and EXOTIC) by established and innovative environmentally friendly low input sustainable methods.

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